174 Species in Genus Bromus

Bromus: A Deep Dive with Ferb Vu

The world of grasses is vast and often overlooked, but to me, it’s a source of endless fascination. Today, I want to delve into a particular genus that has captured my attention: Bromus, more commonly known as brome grasses, cheat grasses, or chess grasses.

These grasses, belonging to the Poaceae family, are found across temperate and cool regions of the globe. They are characterized by their spikelets, which are the basic units of a grass inflorescence. Each spikelet contains multiple florets, small flowers that are typical of grasses.

Why Bromus?

My interest in Bromus stems from their ecological and agricultural significance. Some species are valuable forage grasses, providing nutritious feed for livestock. Others, however, are considered weeds, competing with crops and native plants. This duality makes them a compelling subject for study.

A Diverse Genus

The number of Bromus species is a subject of ongoing debate among taxonomists, with estimates ranging from 100 to 400. However, the current consensus hovers around 174 distinct species.

Here are:

  1. Bromus aegyptiacus Tausch
  2. Bromus alopecuros Poir.
  3. Bromus andringitrensis A.Camus
  4. Bromus anomalus Rupr. ex E.Fourn.
  5. Bromus araucanus Phil.
  6. Bromus arenarius Labill.
  7. Bromus aristatus (K.Koch) Steud.
  8. Bromus arizonicus (Shear) Stebbins
  9. Bromus armenus Boiss.
  10. Bromus arrhenatheroides Baker
  11. Bromus arvensis L.
  12. Bromus attenuatus Swallen
  13. Bromus auleticus Trin. ex Nees
  14. Bromus ayacuchensis Saarela & P.M.Peterson
  15. Bromus benekenii (Lange) Trimen
  16. Bromus berteroanus Colla
  17. Bromus biebersteinii Roem. & Schult.
  18. Bromus bikfayensis A.Camus & Gomb.
  19. Bromus × bolzeanus H.Scholz
  20. Bromus bonariensis Parodi & J.H.Camara
  21. Bromus borianus H.Scholz
  22. Bromus brachyantherus Döll
  23. Bromus brachystachys Hornung
  24. Bromus × brevieri Chass.
  25. Bromus brevis Steud.
  26. Bromus briziformis Fisch. & C.A.Mey.
  27. Bromus bromoideus (Lej.) Crép.
  28. Bromus carinatus Hook. & Arn.
  29. Bromus caroli-henrici Greuter
  30. Bromus catharticus Vahl
  31. Bromus cebadilla Steud.
  32. Bromus ceramicus Ohwi
  33. Bromus chrysopogon Viv.
  34. Bromus ciliatus L.
  35. Bromus coloratus Steud.
  36. Bromus × commutatojaponicus Nyár.
  37. Bromus commutatus Schrad.
  38. Bromus condensatus Hack.
  39. Bromus confinis Nees ex Steud.
  40. Bromus danthoniae Trin.
  41. Bromus densus Swallen
  42. Bromus diandrus Roth
  43. Bromus dolichocarpus Wagnon
  44. Bromus × eburonensis (Nyman) K.Richt.
  45. Bromus elidis H.Scholz
  46. Bromus epilis Keng f.
  47. Bromus erectus Huds.
  48. Bromus exaltatus Bernh.
  49. Bromus fasciculatus C.Presl
  50. Bromus × ferronii Mabille
  51. Bromus firmior (Nees) Stapf
  52. Bromus × fischeri Cugnac & A.Camus
  53. Bromus flexuosus Planchuelo
  54. Bromus formosanus Honda
  55. Bromus frigidus Boiss. & Hausskn.
  56. Bromus frondosus (Shear) Wooton & Standl.
  57. Bromus gedrosianus Pénzes
  58. Bromus gracillimus Bunge
  59. Bromus × granatensis A.Camus
  60. Bromus grandis (Shear) Hitchc.
  61. Bromus grossus Desf. ex DC.
  62. Bromus × guetrotii A.Camus
  63. Bromus gunckelii Matthei
  64. Bromus × hannoverianus K.Richt.
  65. Bromus haussknechtii Boiss.
  66. Bromus himalaicus Stapf
  67. Bromus hordeaceus L.
  68. Bromus husainii P.Agnihotri & R.Yadav
  69. Bromus × husnotii A.Camus
  70. Bromus induratus Hausskn. & Bornm.
  71. Bromus inermis Leyss.
  72. Bromus insignis Buse
  73. Bromus intermedius Guss.
  74. Bromus interruptus (Hack.) Druce
  75. Bromus japonicus Houtt.
  76. Bromus kalmii A.Gray
  77. Bromus kashmirensis P.Agnihotri & R.Yadav
  78. Bromus koeieanus Melderis
  79. Bromus kopetdagensis Drobow
  80. Bromus kozhevnikovii (Prob.) Chepinoga
  81. Bromus × laagei Cugnac & A.Camus
  82. Bromus laevipes Shear
  83. Bromus lanatipes (Shear) Rydb.
  84. Bromus lanatus Kunth
  85. Bromus lanceolatus Roth
  86. Bromus latiglumis (Shear) Hitchc.
  87. Bromus lepidus Holmb.
  88. Bromus leptoclados Nees
  89. Bromus lithobius Trin.
  90. Bromus × litvinovii Roshev. ex Nevski
  91. Bromus macrantherus (Hack. ex Trab.) Henriq.
  92. Bromus macrocladus Boiss.
  93. Bromus madritensis L.
  94. Bromus magnus Keng
  95. Bromus mairei Hack. ex Hand.-Mazz.
  96. Bromus malyschevii (N.V.Vlassova) Chepinoga
  97. Bromus mango É.Desv.
  98. Bromus marginatus Nees ex Steud.
  99. Bromus maritimus (Piper) Hitchc.
  100. Bromus maroccanus Pau & Font Quer
  101. Bromus modestus Renvoize
  102. Bromus moellendorffianus (Asch. & Graebn.) Hayek
  103. Bromus moesiacus Velen.
  104. Bromus moeszii Pénzes
  105. Bromus morrisonensis Honda
  106. Bromus musadoghanii Yıld.
  107. Bromus natalensis Stapf
  108. Bromus nepalensis Melderis
  109. Bromus nottowayanus Fernald
  110. Bromus orcuttianus Vasey
  111. Bromus orientalis Behçet & Yapar
  112. Bromus oxyodon Schrenk
  113. Bromus pacificus Shear
  114. Bromus pannonicus Kumm. & Sendtn.
  115. Bromus parodii Covas & Itria
  116. Bromus paulsenii Hack.
  117. Bromus pectinatus Thunb.
  118. Bromus pellitus Hack.
  119. Bromus picoeuropeanus Acedo & Llamas
  120. Bromus pinetorum Swallen
  121. Bromus pitensis Kunth
  122. Bromus plurinodis Keng
  123. Bromus polyanthus Scribn. ex Shear
  124. Bromus porphyranthos Cope
  125. Bromus porteri (Coult.) Nash
  126. Bromus psammophilus P.M.Sm.
  127. Bromus pseudobrachystachys H.Scholz
  128. Bromus pseudolaevipes Wagnon
  129. Bromus pseudoramosus Keng f. ex L.Liu
  130. Bromus pseudosecalinus P.M.Sm.
  131. Bromus pubescens Muhl. ex Willd.
  132. Bromus pulchellus Fig. & De Not.
  133. Bromus pumilio (Trin.) P.M.Sm.
  134. Bromus pumpellianus Scribn.
  135. Bromus racemosus L.
  136. Bromus ramosus Huds.
  137. Bromus remotiflorus (Steud.) Ohwi
  138. Bromus richardsonii Link
  139. Bromus rigidus Roth
  140. Bromus riparius Rehmann
  141. Bromus × robustus H.Scholz
  142. Bromus × rosettiae A.Camus
  143. Bromus rubens L.
  144. Bromus salangensis Naderi
  145. Bromus sclerophyllus Boiss.
  146. Bromus scoparius L.
  147. Bromus scopulorum Chase
  148. Bromus secalinus L.
  149. Bromus segetum Kunth
  150. Bromus × segoviensis A.Camus
  151. Bromus setifolius J.Presl
  152. Bromus sewerzowii Regel
  153. Bromus sinensis Keng f.
  154. Bromus sipyleus Boiss.
  155. Bromus sitchensis Trin.
  156. Bromus speciosus Nees
  157. Bromus squarrosus L.
  158. Bromus staintonii Melderis
  159. Bromus stenostachyus Boiss.
  160. Bromus sterilis L.
  161. Bromus striatus Hitchc.
  162. Bromus suksdorfii Vasey
  163. Bromus sundaicus Ohwi
  164. Bromus syriacus Boiss. & C.I.Blanche
  165. Bromus tectorum L.
  166. Bromus texensis (Shear) Hitchc.
  167. Bromus timorensis Veldkamp
  168. Bromus tomentellus Boiss.
  169. Bromus tomentosus Trin.
  170. Bromus tunicatus Phil.
  171. Bromus tyttholepis (Nevski) Nevski
  172. Bromus variegatus M.Bieb.
  173. Bromus villosissimus Hitchc.
  174. Bromus vulgaris (Hook.) Shear

Identification Challenges

Identifying Bromus species can be quite challenging, even for experienced botanists. The differences between some species are subtle and can be easily overlooked. Key features to examine include the size and shape of the spikelets, the presence and length of awns (bristle-like appendages on the florets), and the hairiness of the leaves and sheaths.

Ecological Impact

Bromus species play a significant role in their respective ecosystems. Some, like smooth brome, are beneficial, providing forage and stabilizing soils. Others, like cheatgrass, can have detrimental effects. Cheatgrass, with its ability to thrive in disturbed areas and its flammable nature, has increased the frequency and intensity of wildfires in the western United States, threatening native plant communities and altering wildlife habitats.

Economic Importance

From an economic standpoint, Bromus species have both positive and negative impacts. Forage grasses like smooth brome contribute to livestock production, while weedy species like cheatgrass can reduce crop yields and increase weed control costs.

Future Research

There’s still much to learn about Bromus. Further research is needed to better understand the ecological roles of different species, their potential for invasiveness, and their response to climate change. This knowledge will be crucial for developing effective management strategies and ensuring the sustainability of our ecosystems and agricultural practices.

I hope this exploration of the Bromus genus has shed light on its complexity and importance. It’s a reminder that even seemingly mundane grasses can have a profound impact on our world. As I continue my botanical journey, I look forward to uncovering more secrets hidden within the world of Bromus.

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